Guthrie, F., mathematician and botanical collaborator of Harry Bolus on the taxonomy of Erica for Flora Capensis. He assembled and excellent herbarium, which is now incorporated in the Bolus Herbarium.
WCM Protologue Reference:
Pillans, Trans. Roy. Soc. South Africa 16: 371 (1928)
Vegetative parts: plants tufted, 0.2–0.6 m tall, basal diameter 0.1–0.3 m, crown diameter 0.3–0.4 m, compact, without spreading rhizomes or stolons. Fertile culms unbranched, smooth, olivaceous, 0.2–0.6 m tall, diameter at base 1–2 mm, diameter at apex 0.6–1.6 mm. Clusters of sterile branches at the nodes of the fertile branches present. Sterile branches present post flowering. Sheaths closely convoluted, 30–60 mm long, dark brown at base,red brown with speckles, apical margins broadly chartaceous and soon decaying, hyaline shoulders absent, apex, excluding the membranous lobes, acute. Sheath-mucro absent or penicellate, straight and erect, 0–3 mm long. Sterile branch leaf-sheath 2–3 mm long, hyaline shoulder 3–5 mm long, lacerated, leaf-blade 4–7 mm long; leaf blade 0.3 mm wide. Males: male inflorescence with 21–50 spikelets, paniculate, 40–120 mm long, 25–50 mm wide. Male inflorescence spathes persistent, upper margins lacerated and largely decayed at anthesis, shorter than spikelets, chartaceous. Male flowers in spikelets. Male spikelets pendulous on flexible pedicels, oblong, apex rounded, 10–20 mm, 4–6 mm, with 10–32 flowers, pedicels shorter than the spikelets, flattened. Male bracts taller than flowers, 5–8.5 mm long, oblong, acute, hyaline (sometimes a broad hyaline margin, sometimes not) or chartaceous or cartilaginous (at least in middle of bract), bract upper margin like body of bract, bract awn minute or absent, brick red. Male flower 2.8–4 mm long. Male tepals sepals more rigid than petals, chartaceous or cartilaginous, petals chartaceous, oblong, inner and outer tepals the same length. Male outer lateral tepals keeled or winged, glabrous. Anthers 1.5–2 mm long, included in the flowers. Pistillode absent. Females: female inflorescence linear or paniculate or racemose, 30–120 mm long, 10–30 mm wide, with 2–5 spikelets or 6–10 spikelets. Female spathes longer than the spikelets, not obscuring spikelet, persistent, chartaceous. Female spikelet pedicel simple or pedicel sparsely branching, oblong, apex rounded, 15–30 mm long, with 10–30 flowers, with 0 sterile bracts. Female bracts at least twice as long as flowers, 10–17 mm long, oblong, acute, chartaceous or cartilaginous, apical margin like rest of bract, same as body of bract, awn minute or absent. Female flower 4–5 mm long, without a fleshy pedicel. Female tepals cartilaginous, glabrous and smooth, at least as tall as nut, wrapped around it and obscuring the surface, midrib flush with the tepal body, apices acute, inner whorl shorter than outer. Female outer lateral tepals keeled or winged, wings entire, wings lateral sepals decurrent, without grooves, as wide as body or wider than body, 4–5 mm long. Female odd outer tepal oblong, 3.5–4.5 mm long. Female inner tepals oblong, 3 mm long. Staminodes absent. Styles plumose, white, 1. Ovary with 1 locule, indehiscent. Fruit: indehiscent fruits with a softer ovary wall. Perianth presence on fruits persistent, winged. Shape of winged dispersal unit oblong, narrower than long. Nut apex smooth, without a distinct cap, wall smooth (rough in upper half from tuberculate cells), 2.1–3 mm long, 1.2–1.8 mm diam., shape in side view elliptical or obovate or oblong, shape in top view elliptical in cross section, brown. Elaiosome absent.